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This is in response to the question raised - what is the place or ranking of systematics in paleontology? The way I see it there are two ways of looking at this. The first it to view systematics as a means of understanding phylogenetic relationships among organisms, establishing evolutionary pathways and reconstructing the course of evolution through time. As an end in itself, I would say that this aspect of systematics would rank equally in value with other branches of paleontology, such as paleoecology, biostratigraphy, etc. Clearly, understanding the history of life on earth is one of our primary goals. The second aspect is that systematics is the means by which we define taxa, establish criteria by which we can indentify particular taxa and the methodological approach we use do this. Identifications of particular taxa at particular points in time and space are the primary bits of data for most paleontological applications such as biostratigraphy, paleobiogeography, etc. It stands to reason, therefore, that identification of taxa is the single most important part of paleontological data aquisition. Misidentifications and identifications of poorly defined taxa are bad data in the same way as an incorrect radiometric date or geochemical analysis. Systematics is the "instrumation" by which we aquire taxonomic identifications and is therefore fundamental to any paleontologic application that uses occurrences of taxa as the data points. As an example of how systematic approach can affect the outcome of a paleontological study, I was involved, a number of years ago (and continue to be involved in) occurrences of graptolites across the Late Ordovician extinction event. Using the traditional means of defining diplograptid genera on the basis of thecal form, some seven or eight genera were known to span this boundary. Employing different criteria to define genera (astogenetic growth patterns + thecal form) combined with a phylogenetic systematic approach, it became apparent that only one or two genera span this boundary and many others become extinct. In addition, the newly defined are much more biostratigraphically restricted than the "old" ones (hence, more useful). In this case, a change in systematic approach resulted not only in (hopefully) better taxonomic definitions (ie. better data), but also in a completely new view of one of the major events in the history of life. We cannot abandon the pursuit of systematics, either in terms of improvement in methodology or in its practice in particular taxonomic groups, any more than chemists can give up on developing, improving, maintaining and using their analytical instruments. Mike Melchin, Chair Geology Dept., St. F.X. University P.O. Box 5000, Antigonish, N.S., Can B2G 2W5 902-867-5177, FAX - 902-867-5153 mmelchin@juliet.stfx.ca
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