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paleonet Confuciusornis bereft after all etc.



Ah - there you are.

Well done on University Challenge in beating the Eden
project.  I couldn't have bettered the captain's
performance, but if you'd manufactured a place for me
alongside him on your team you'd have reached the
finals.

Anyway...

Perhaps the most fascinating nugget to fall out on
opening "Mesozoic Birds" (presumably the add-on title
"Above the Heads of Dinosaurs" is a jibe at what they
see as the "Old Guard") is a mention by Zhou & Hou on
p165 that Confuciusornis does in fact NOT have
uncinate processes.

One can only imagine co-editor Luis Chiappe's reaction
to this, having co-authored the 1999 monograph on
Confuciusornis containing two photos of the rib hooks.
 Somewhere in the book the editors make a point of
stating that Zhou & Hou's material was not the same as
that used for the monograph, but this patch doesn't
quite heal the wound.  Although they might seem a tiny
skeletal detail, UPs are the most informative part of
any Cretaceous mani. because it is, as far as I can
tell, the ONLY reliable character; in the baffling
chaos of early avian phylogeny, beset by two separate
attractors, one evolving multiple lineages towards
enhanced flight ability, and the other converging them
towards flightlessness, if we are lucky enough to find
some reliable anchor it can help fix our views on
other features.  UPs provide the edge pieces bordering
our 1000-piece jigsaw.  (I won't be repeating why
today.)

Their absence from Conf. will be annoying to the
palaeontological establishment for so many reasons. 
Not only has Storrs Olson blasted away about their
absence in the past, but I too have been saying it for
years (it was edited out of my review of Paul 2002 -
THES 26.4.02 p26).  It's a mark of a good theory when
it throws doubt on accepted observations and that
doubt is then justified, but publicity is required for
the scientific process to benefit from this, and it
seems I can't leave this to you.

Chiappe, Norell, and all their confederates at AMNH;
Ji Shu'an & Ji Qiang; Paul; Milner and her associates
at NHM, and all the others who have stuck UPs on
Confuciusornis down the years despite my advice to the
contrary, have to face the fact that it doesn't matter
how many real examples you've breathed over, you can't
match someone with a few good photos and a working
thinking cap on.  The couple of supposed images of UPs
in the Conf. monograph, while not very convincing are
not quite as unbelievable as I at first thought. 
However Olson was right: it is the statistics of the
evidence that is absolutely undeniable.  If those two
scrappy examples are the best to be found from a taxon
orders of magnitude better represented than most
Cretaceous dino-birds, the chance the 'UP's were real
was absolutely infeasible.  Even if the billion to one
odds against were actually right, it would not be
scientifically responsible to accept them. It's no
good saying "they're poorly preserved features, and
few ovis and droms preserve them", any realistic
statistical model will still send the message through
loud and clear, especially when near perfect
side-preserved examples like the one at the bottom of
page 86 in "DA" don't show them.

Mark my words, UPs of a certain form evolved only once
and were completely lost in no significant radiations
over some 150 million years.  If true, that's
incredibly useful, and it would be the greatest folly
to resist the idea just because you didn't think of
it.  The few "enants" left that still have them aren't
enants. Their invention by flying droms reflects the
truly bizarre requirements of their lifestyle; serious
thought needs to be given as to what extent
non-uncinated fliers could breathe and flap at
different frequencies.

Other parts of "Mesozoic Birds" are interesting too. 
For example seeing all the alvarezsaur pieces gathered
together showed me that the northern hemisphere types
are descended from flightless troodonts, but the S Am.
ones aren't, though they might or might not be
descended from flying ones via droms.  If the S Am.
ones are flightless droms as I suspect, both N & S
alvzs. are birds, but they're still poly- (at least
di-) phyletic.

I was interested to see George Olshevsky indexed the
book, and that neoflightlessness theories are given
some mention eg on p22, though that page and many many
others also ship unscientific drivel on an industrial
scale - for example when claiming the only evidence
comes from simplistic cladism, and that cladograms
test anything.  Your senior supplier of cladogram
programs has told you they will output garbage when
used on data with as much inherent parallelism as
found with dino-birds, in: 
Felsenstein, J. 1978. Cases in which parsimony or
compatibility methods will be positively misleading.
Systematic Zoology 27: 401-410 (Also reprinted as pp.
663-674 in Conceptual Issues in Evolutionary Biology,
An Anthology, ed. Elliott Sober, MIT Press, Cambridge,
Massachusetts, 1984), 
a paper which the SCI shows has been referenced by
almost every one in the world except dino-bird
palaeontologists.  In what other industry could
salaries be drawn for so many years on work based on
such a flouting of the "GIGO" dictum?  Nigel Calder's
recent comments on abuses of the academic system could
be made for you! 

JJ


		
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