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In a message dated 4/13/2005 12:58:22 P.M. Pacific Standard Time,
tijawi@yahoo.com writes:
>>A cladogram can indeed tell you the order in which individual characters
arose. For example, a cladogram of the Dinosauria tells us that theropods
inherited obligate bipedalism, and then went on to evolve (in the following
order): furcula ("wishbone"); feathers; semilunate carpal ("swivel-wrist' -
essential for execution of the flight stroke); elongated forelimbs; asymmetric
vanes; and reversed hallux. This sequence of character acquisition contradicts
George's own hypothesis, so he has a vested interest in distrusting what the
cladogram is telling him.<<
No, it cannot. This is an assumption built into the cladogram by choice of
outgroup. You choose a bipedal form as the outgroup, and sure, your cladogram
will "prove" that the ancestor of the clade is a biped(!). The problem with
most dinosaur cladograms is that they have many character polarities backward
ab initio.
Semilunate carpal evolved at least twice in Maniraptora (segnosaurs and
deinonychosaurs independently: Alxasaurus had no semilunate carpal, eg). Reversed
hallux evolved very early in theropods and occurs in all theropods at or
above the ceratosaur level--look at theropod foot anatomy, fer chrissake, or at
theropod footprints from the Triassic. It is not "essential" for the
execution of the flight stroke, it is an adaptation to make the flight stroke more
efficient. The flight stroke comes first, then the improvements, not vice versa.
Feathers? We know nothing about the evolution of feathers from any
cladogram. But do you think they appeared fully formed by magic in Archaeopteryx?
Bipedality. I claim that many dinosaurs were bipedal because the forelimbs
of their volant ancestors had become too winglike to be used for walking by
their ground-dwelling descedants. Current models of dinosaur evolution provide
NO reason for the appearance of bipedality in dinosaurs except the
tautological one: they somehow made dinosaurs "better."
And so on.
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