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In a message dated 4/13/2005 12:58:22 P.M. Pacific Standard Time, tijawi@yahoo.com writes: >>A cladogram can indeed tell you the order in which individual characters arose. For example, a cladogram of the Dinosauria tells us that theropods inherited obligate bipedalism, and then went on to evolve (in the following order): furcula ("wishbone"); feathers; semilunate carpal ("swivel-wrist' - essential for execution of the flight stroke); elongated forelimbs; asymmetric vanes; and reversed hallux. This sequence of character acquisition contradicts George's own hypothesis, so he has a vested interest in distrusting what the cladogram is telling him.<< No, it cannot. This is an assumption built into the cladogram by choice of outgroup. You choose a bipedal form as the outgroup, and sure, your cladogram will "prove" that the ancestor of the clade is a biped(!). The problem with most dinosaur cladograms is that they have many character polarities backward ab initio. Semilunate carpal evolved at least twice in Maniraptora (segnosaurs and deinonychosaurs independently: Alxasaurus had no semilunate carpal, eg). Reversed hallux evolved very early in theropods and occurs in all theropods at or above the ceratosaur level--look at theropod foot anatomy, fer chrissake, or at theropod footprints from the Triassic. It is not "essential" for the execution of the flight stroke, it is an adaptation to make the flight stroke more efficient. The flight stroke comes first, then the improvements, not vice versa. Feathers? We know nothing about the evolution of feathers from any cladogram. But do you think they appeared fully formed by magic in Archaeopteryx? Bipedality. I claim that many dinosaurs were bipedal because the forelimbs of their volant ancestors had become too winglike to be used for walking by their ground-dwelling descedants. Current models of dinosaur evolution provide NO reason for the appearance of bipedality in dinosaurs except the tautological one: they somehow made dinosaurs "better." And so on.
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