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Re: paleonet heterogenous taxa and ICZN regulations



Ken

 

There is for example a publication of 1901, in which a new genus is

described, consisting of two species. The first issue concerns the type

species (I'll call it here 'Species 1'). In its description, twelve

specimens are mentioned and each briefly described. These specimens are

however a heterogenous assembly of two different species with clearly

different morphological traits.

 

This is a common problem in all taxonomic groups and is dealt with successfully on a daily basis.

 

My colleague therefore states that its

genus name has to be deleted because its original definition is

incorrect. It should thus be either a nomen dubium or a synonym (if

assumed that the type belongs to a previously existing genus-something I

disagree with).

 

Your colleague does not know what he/she is talking about.

 

I think it doesn't add up. Isn't the name of the species

fixed by its holotype, prompting the need to revise the species by

excluding the original paratypes that are something else?

 

Yes. The specific name is attached in perpetuity to the holotype specimen, or to the lectotype if there is initially no specimen identified as a holotype. It does not matter what happens to that specimen later. Whether it is synonymised with another species, assigned to another genus, found to be part of a larger organism, is immaterial. That specific name belongs with that specimen.

 

Such a case is not explicitly stated by ICZN as far as I see, but I always assumed it

works this way. Am I right?

 

Yes, you are right. The ICZN is not explicit on many possible scenarios, but the correct procedure is implicit in the framing of the rules.

 

To make things more complicated, another

species is described from the same genus. However, this 'Species 2' is

not congeneric with 'Species 1', in the sense that the holotypes of

Species 1 and 2 belong to different genera with different morphological

traits.

 

This does not complicate things much at all. Again, it is a scenario that occurs every day. Not all genera now recognised are based on newly discovered specimens of previously unknown species. Palaeontologists have been subdividing genera since Linnaeus fell of his perch.

 

My colleague would say, in a similar fashion, that the genus

name is redundant, because its original definition is incorrect.

 

Your colleague hasn’t been taking his/her tablets! Whether someone’s work is incorrect is a matter of opinion. Someone may in future come along and decide that your fish specimens all belong to the same species and that you misconstrued metamorphosis for interspecific difference.

 

Again, ICZN doesn't explicitly address an issue [such] as this, but again I'd assume,

as with the species example, that its name is fixed by the type species

and that the genus needs to be revised. Do I see it right?

 

Yes. The genus is attached to the type species, similar to the way the species is attached to the holotype. Even if someone synonymises that species with another, the generic name adheres to that type species. This is because, the synonymy is simply an opinion and some later worker may come along and decide that the synonymy is incorrect!

 

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Dr John R. Laurie

Eastern and Onshore Petroleum

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